Note: you can download pdf copies of my articles, particularly those where I am the first author, if they will exclusively be used for research.

Peer-reviewed publications


[8] Fajardo, A., and Velázquez, E. (2021) Fine-scale spatial associations between functional traits and tree growth. Oikos, accepted (Funded by Fondecyt 1160329, 1190900)

[7] Fajardo, A., Llancabure, J.C. and Moreno, P.C. (2021) Assessing forest degradation using multivariate and machine learning methods in the Patagonian temperate rainforest. Ecological Applications, accepted (Funded by FIBN 2016/003)

[6] Camarero, J.J., Gazol, A., Linares, J.C., Fajardo, A., Colangelo, M., Valeriano, C., et al. (2021) Differences in temperature sensitivity and drought recovery between natural stands and plantations of conifers are species-specific. Science of the Total Environment 796: 148930 (Funded by Fondecyt 1190900)

[5] Camarero, J.J., Gazol, A., Sánchez-Salguero, R., Fajardo, A., McIntire, E.J.B., Liang, E. (2021) Tree growth and treelike responses to temperature: Different questions and concepts. Global Change Biology 27(17): e13-e14

[4] Fajardo, A., and Piper, F.I. (2021) How to cope with drought and not die trying: drought acclimation across tree species with contrasting niche breadth. Functional Ecology 35(9): 1903-1913 (Funded by Fondecyt 1160329, 1190900)

[3] Nuske, S.J., Fajardo, A., Nuñez, M.A., Pauchard, A., Wardle, D.A., Nilsson, M.-C., Kardol, P., Smith, J.E., Peltzer, D.A., Moyano, J., and Gundale, M.J. (2021) Soil biotic and abiotic effects on seedling growth exhibit context dependent interactions: evidence from a multi-country experiment on Pinus contorta invasion. New Phytologist 232(1): 303-317 (Funded by Fondecyt 1190900)

[2] Koçillari, L., Olson, M.E., Suweis, S., Rocha, R.P., Lovison, A., Cardin, F., Dawson, T.E., Echeverría, A., Fajardo, A., et al. (2021) The widened pipe model of plant hydraulic evolution. Proceedings of the National Academy of Science of the USA 118(22): e2100314118 (Funded by Fondecyt 1190900)

[1] Camarero, J.J., Gazol, A., Sánchez-Salguero, R., Fajardo, A., McIntire, E.J.B., Gutiérrez, E., et al. (2021) Global fading of the temperature-growth coupling at alpine and polar treelines. Global Change Biology 27(9): 1879-1889 (Funded by Fondecyt 1160329)


[4] Fajardo, A., Mora, J.P., and Robert, E. (2020) Corner’s rules pass the test of time: little effect of phenology on leaf-shoot and other scaling relationships. Annals of Botany 126(7): 1129-1139, doing:10.1093/aob/mcaa124 (Funded by Fondecyt 1160329)

[3] Olson, M.E., Rosell, J.A., Martínez-Pérez, C., León-Gómez, C., Fajardo, A., Isnard, S., Cervantes-Alcayde, M.A., Echeverría, A., Figueroa-Abundiz, V.A., Segovia-Rivas, A., Trueba, S., and Vásquez-Segovia, K. (2020) Xylem vessel diameter-shoot length scaling: ecological significance of porosity types and other traits. Ecological Monographs 90(3): e01410, doi:10.1002/ecm.1410 (Funded by Fondecyt 1160329)

[2] García-Cervigón, A.I., Fajardo, A., Caetano-Sánchez, C., Camarero, J.J., and Olano, J.M. (2020) Xylem anatomy needs to change, so that conductivity can stay the same: xylem adjustments across elevation and latitude in Nothofagus pumilio. Annals of Botany 125(7): 1101-1112 (Funded by Fondecyt 1160329)

[1] Fajardo, A., Martínez-Pérez, C., Cervantes-Alcayde, M.A., and Olson, M.E. (2020) Stem length, not climate, controls vessel diameter in two species across a sharp precipitation gradient. New Phytologist 225(6): 2347-235, doi: 10.1111/nph.16287 (Funded by Fondecyt 1160329)


[8] Fajardo, A., and Piper, F.I. (2019) Cluster-root bearing Proteaceae species show a competitive advantage over non-cluster root-bearing species. Annals of Botany 124(6): 1121-1131

[7] Piper, F.I., Fajardo, A., Baeza, G., and Cavieres, L.A. (2019) The association between a nurse cushion plant and a cluster-root bearing tree species alters the plant community structure. Journal of Ecology, 107(5): 2182-2196

[6] Piper, F.I., Gundale, M.J., Fuenzalida, T., and Fajardo, A. (2019) Herbivore resistance in congeneric and sympatric Nothofagus species is not related to leaf habit. American Journal of Botany 106(6): 788-797

[5] Fajardo, A., Gazol, A., Mayr, C., and Camarero, J.J. (2019) Recent decadal growth reverts warming-triggered growth enhancement in contrasting climates in the southern Andes tree line. Journal of Biogeography 46(7): 1367-1379 (Funded by Fondecyt 1160329)

[4] Fajardo, A., and Siefert, A. (2019) The interplay among intraspecific leaf trait variation, niche breadth and species abundance along light and soil nutrient gradients. Oikos 128(6): 881-891 (Funded by Fondecyt 1160329)

[3] Fajardo, A., McIntire, E.J.B., and Olson, M.E. (2019) When short stature is an asset in trees. Trends in Ecology and Evolution 34(3): 193-199 (Funded by Fondecyt 1160329)

[2] Piper, F.I., Hoch, G., and Fajardo, A. (2019) Revisiting the relative growth rate hypothesis for gymnosperm and angiosperm species co-occurring in southern Chile. American Journal of Botany 106(1): 101-112

[1] Davis, K.T., Callaway, R.M., Fajardo, A., Pauchard, A., Nuñez, M.A., Brooker, R.W., . . . Gundale, M.J. (2019) Severity of impacts of an introduced species corresponds with regional eco-evolutionary experience. Ecography 42(1): 12-22


[7] Craine, J.M., Elmore, A.J., Wang, L., Aranibar, J., Bauters, M., Boeckx, P., Crowley, B., Dawes, M.A., Delzon, F., Fajardo, A., . . . Zmudczyńska-Skarbek, K. (2018) Isotopic evidence for oligotrophication of terrestrial ecosystems. Nature Ecology & Evolution, in press

[6] Olson, M.E., Soriano, D., Rosell, J.A., Anfodillo, T., León-Gómez, C., Camarero, J.J., Castorena, M., Dawson, T., Donoghue, M.J., Echeverría, A., Edwards, E.J., Espinosa, C.T., Fajardo, A., Gazol, A., Isnard, S., Lima, R., Marcati, C.R. and Méndez-Alonzo, R. 2018. Plant height and hydraulic vulnerability to drought and cold. Proceedings of the National Academy of Science of the USA 115(29): 7551-7556 (Funded by: Fondecyt 1120171)

[5] García-Cervigón, A., Olano, J.M., von Arx, G., and Fajardo, A. 2018. Xylem adjusts to maintain efficiency across a steep precipitation gradient in two coexistent generalist species. Annals of Botany 122(3): 461-472 (Funded by: Fondecyt 1160329)

[4] Camarero, J.J., Gazol, A., Sangüesa-Barreda, G. and Fajardo, A.2018. Coupled climate-forest growth shifts in the Chilean Patagonia are decoupled from trends in water-use efficiency. Agricultural and Forest Meteorology 259(1): 222-231 (Funded by: Fondecyt 1160329)

[3] Fajardo, A., and Siefert, A. 2018. Intraspecific trait variation and the leaf economics spectrum across resource gradients and levels of organizationEcology 99(5): 1024-1030 (Funded by: Fondecyt 1120171)

[2] Fajardo, A. 2018. Insights into intraspecific wood density variation and its relationship with growth, height and elevation in a treeline speciesPlant Biology20(3): 456-464 (Funded by: Fondecyt 1120171)

[1] Fajardo, A., and Gundale, M.J. 2018. Canopy cover type, and not fine-scale resource availability, explains native and exotic species richness in a landscape affected by anthropogenic fires and posterior land-use changeBiological Invasions 20(2): 385-398 (Funded by: Fortalecimiento SS-2008-10 & Fondecyt 1120171)


[4] Piper, F.I., Fajardo, A., and Hoch, G. 2017. Single-provenance mature conifers show higher non-structural carbohydrate storage and reduced growth in a drier location. Tree Physiology 37(8): 1001-1010.

[3] Camarero, J.J., and Fajardo, A. 2017. Poor acclimation to current drier climate of the long-lived tree species Fitzroya cupressoides in the temperate rainforest of southern Chile. Agricultural and Forest Meteorology 239(1): 141-150

[2] Fajardo, A., and Piper, F.I. 2017. An assessment of carbon and nutrient limitations in the formation of the southern Andes treelineJournal of Ecology 105(2): 517-527 (Funded by: Fondecyt 1120171)

[1] Mayor, J.R., Sanders, N.J., Classen, A.T., Bardgett, R.D., Clément, J.-C., Fajardo, A., Lavorel, S., Sundqvist, M.K., Bhan, M., Chisholm, C., Cieraad, E., Gedalof, G., Grigulis, K., Kudo, G., Oberski, D., and Wardle, D.A. 2017. Elevation alters ecosystem properties across temperate treelines globallyNature 542(7639): 91-95 (Funded by: Fondecyt 1120171)


[11] Fajardo, A., and Siefert, A. 2016. Temperate rain forest species partition fine-scale gradients in light availability based on their leaf mass per area (LMA). Annals of Botany 118(7): 1307-1315 (Funded by: Fondecyt 1120171)

[10] Piper, F.I., and Fajardo, A. 2016. Carbon dynamics of Acer pseudoplatanus seedlings under drought and complete darkness. Tree Physiology 36(11): 1400-1408

[9] McIntire, E.J.B., Piper, F.I., and Fajardo, A. 2016. Wind and light exposure, more than elevation-related temperature, limit treeline seedling abundance on three continents. Journal of Ecology, 104(5): 1379-1390

[8] Gundale, M.J., Almeida, J.P., Wallander, H., Wardle, D.A., Kardol, P., Nilsson, M.-C., Fajardo, A., Pauchard, A., Peltzer, D.A., Ruotsalainen, S., Mason, B., and Rosenstock, N. 2016. Differences in endophyte communities of introduced trees depend on the phylogenetic relatedness of the receiving forest. Journal of Ecology, 104(5): 1219-1232

[7] Bravo-Monasterio, P., Pauchard, A., and Fajardo, A. 2016. Pinus contorta invasion into treeless steppe reduces species richness and alters species traits of the local community. Biological Invasions, 18(7): 1883-1894 (Funded by: Fondecyt 1120171)

[6] Fajardo, A. 2016. Are trait-scaling relationships invariant across contrasting elevations in the widely distributed treeline species Nothofagus pumilio? American Journal of Botany, 103(5): 821-829 (Funded by: Fondecyt 1120171)

[5] Piper, F.I., Viñegla, B., Linares, J.C., Camarero, J.J., Cavieres, L.A., and Fajardo, A. 2016. Mediterranean and temperate treelines are controlled by different environmental drivers. Journal of Ecology 104(3): 691-702

[4] Fajardo, A. 2016. Wood density is a poor predictor of competitive ability among individuals of the same species. Forest Ecology and Management372(1): 217-225 (Funded by: Fondecyt 1120171)

[3] Fajardo, A., and Siefert, A. 2016. Phenological variation of leaf functional traits within species. Oecologia 180(4): 951-960 (Funded by: Fondecyt 1120171)

[2] Renard, S., McIntire, E.J.B., and Fajardo, A.  2016. Winter conditions—not summer temperature—influence establishment of seedlings at white spruce alpine treeline in Eastern Quebec. Journal of Vegetation Science 27(1): 29-39

[1] Fajardo, A., Torres-Díaz, C., and Till-Bottraud, I. 2016. Disturbance and density-dependent processes (competition and facilitation) influence the fine-scale genetic structure of a tree species population. Annals of Botany 117(1): 67-77 (Funded by: Fondecyt 1120171 & Ecos-Conicyt C12B01)


[4] Siefert, A., Violle, C., Chalmandrier, L., Albert, C.H., Taudiere, A., Fajardo, A., Aarssen, L.W. et al. 2015. A global meta-analysis of the relative extent of intraspecific trait variation in plant communities. Ecology Letters 18(12): 1406-1419 (Funded by: Fondecyt 1120171)

[3] Fajardo, A., and Gundale, M.J. 2015. Combined effects of anthropogenic fires and land-use change on soil properties and processes in Patagonia, Chile. Forest Ecology and Management 357(1): 60-67 (Funded by: Fortalecimiento SS-2008-10)

[2] Piper, F.I., Gundale, M.J., and Fajardo, A. 2015. Extreme defoliation reduces tree growth but not C and N storages in a winter deciduous species. Annals of Botany 115(7): 1093-1103 (Funded by: DID S-2010-67)

[1] Fajardo, A. and Piper, F.I. 2015. High foliar nutrient concentrations and resorption efficiency in Embothrium coccineum (Proteaceae) in southern ChileAmerican Journal of Botany 102(2): 208-216 (Funded by: Fondecyt 1120171)


[3] Piper, F.I. and Fajardo, A. 2014. Foliar habit, tolerance to herbivory, and their link to carbon and nitrogen storage. Journal of Ecology, 102(5): 1101-1111 (Funded by: DID S-2010-67)

[2] Fajardo, A. and Piper, F.I. 2014. An experimental approach to explain the Southern Andes elevational treeline. American Journal of Botany 101(5): 788-795 (Funded by: Fondecyt 3070050)

[1] McIntire, E.J.B. and Fajardo, A. 2014. Facilitation as a ubiquitous driver of biodiversity (Tansley Review)New Phytologist 201(2): 403-416


[4] Piper, F.I., Baeza, G., Zúñiga-Feest, A. and Fajardo, A. 2013. Soil nitrogen, and not phosphorus, promotes cluster root formation in a South American Proteaceae. American Journal of Botany 100(12): 2328-2338 (Funded by: DID S-2012-55). Media report about our article: 

[3] Adler, P.B., Fajardo, A., Kleinhesselink, A.R., and Kraft, N.J.B. 2013. Trait-based tests of coexistence mechanisms. Ecology Letters 16(10): 1294-1306(Funded by: Fondecyt 1120171)

[2] Fajardo, A., Piper, F.I., and Hoch, G. 2013. Similar variation in carbon storage between deciduous and evergreen treeline species across elevational gradients. Annals of Botany 112(3): 623-631

[1] Piper, F.I., Fajardo, A. and Cavieres, L.A. 2013. Simulated warming does not impair seedling survival and growth of Nothofagus pumilio in the Southern Andes. Perspectives in Plant Ecology, Evolution and Systematics 15(2): 97-105


[4] Till-Bottraud, I., Fajardo, A. and Rioux, D. 2012. Multi-stemmed trees of Nothofagus pumilio second-growth forests in Patagonia are formed by highly related individuals. Annals of Botany 110(4): 905-913

[3] Fajardo, A., Piper, F.I., Pfund, L., Korner, C. and Hoch, G. 2012. Variation of mobile carbon reserves in trees at the alpine treeline ecotone is under environmental control. New Phytologist 195(4): 794-802

[2] Bravo, P., Penailillo, P. and Fajardo, A. 2012. New antecedents on the distribution and ecology of Botrychium dusenii (H. Christ.) Alston (Ophioglossaceae) associated with anthropogenic environments in the Coyhaique province, Aysen Region, Chile. Gayana Botanica 69(1): 208-211

[1] Fajardo, A. and McIntire, E.J.B. 2012. Reversal of multi-century tree growth improvements and loss of synchrony at mountain treelines point to changes in key driversJournal of Ecology 100(3): 782-794


[6] Gundale, M.J., Fajardo, A., Lucas, R.W., Nilsson, M.-C. and Wardle, D.A. 2011. Resource heterogeneity does not explain the productivity-diversity relationship across a boreal island fertility gradientEcography 34(5): 887-896

[5] Piper, F.I. and Fajardo, A. 2011. No evidence of carbon limitation with tree age and height in Nothofagus pumilio under Mediterranean and temperate climate conditions. Annals of Botany 108(5): 907-917. Reviewed in Faculty of 1000 as a “must read” paper.

[4] McIntire, E.J.B. and Fajardo, A. 2011. Facilitation within species: a possible origin of group selected superorganisms. The American Naturalist 178(1): 88-97

[3] Fajardo, A. and McIntire, E.J.B. 2011. Under strong niche overlap conspecifics do not compete but help each other to survive: facilitation at the intraspecific levelJournal of Ecology 99(2): 642-650

[2] Fajardo, A., Piper, F.I. and Cavieres, L.A. 2011. Distinguishing local from global climate influences in the variation of carbon status with altitude in a tree line speciesGlobal Ecology and Biogeography 20(2): 307-318

[1] Fajardo, A., and Piper, F.I. 2011. Intraspecific trait variation and covariation in a widespread tree species (Nothofagus pumilio) in southern ChileNew Phytologist 189(1): 259-271

2010 and before

[11] Fajardo, A., and McIntire, E.J.B. 2010. Merged trees in second-growth, fire origin forests in Patagonia, Chile: positive spatial association patterns and their ecological implications. American Journal of Botany 79(9): 1424-1430

[10] Fajardo, A., and González, M.E. 2009.  Replacement patterns and species coexistence in an Andean AraucariaNothofagus forestJournal of Vegetation Science 20(6): 1176-1190

[9]  McIntire, E.J.B. and Fajardo, A. 2009.  Beyond description: the active and effective way to infer processes from spatial patternsEcology 90(1): 46-56

[8] Fajardo, A., Quiroz, C., and Cavieres, L.A.  2008.  Distinguishing colonization modes from spatial structures in populations of the cushion plant Azorella madreporica in the high-Andes of central ChileAustral Ecology 33(6): 703-712

[7] Fajardo, A., Quiroz, C., and Cavieres, L.A. 2008.  Spatial patterns in cushion-dominated plant communities of the high-Andes of central Chile: how frequent are positive associations?  Journal of Vegetation Science 19(1): 87-96

[6] Fajardo, A., and McIntire, E.J.B. 2007.  Distinguishing microsite and competition processes in tree growth dynamics: an a prioriThe American Naturalist 169(5): 647-661

[5] Fajardo, A., Graham, J.M., Goodburn, J.M., and Fiedler, C.E. 2007.  Ten-year responses of ponderosa pine growth, vigor, and recruitment to restoration treatments in the Bitterroot Mountains, Montana, USAForest Ecology and Management 243(1): 50-60

[4] Fajardo, A., Goodburn, J., and J. Graham. 2006.  Spatial patterns of ponderosa pine and Douglas fir saplings following selection harvests in western Montana, USAForest Ecology and Management 223(1-3): 255-266

[3] Fajardo, A., and Alaback, P. 2005.  Effects of natural and human disturbances on the dynamics and spatial structure of Nothofagus glauca (Phil.) Krasser forests in South-central ChileJournal of Biogeography 32(10): 1811-1825

[2] Cavieres, L.A., and Fajardo, A. 2005.  Browsing by large herbivores on Nothofagus pumilio forest gaps in Tierra del Fuego, ChileForest Ecology and Management 204(2-3): 237-248

[1] Fajardo, A., and de Graaf, R. 2004.  Tree dynamics in canopy gaps in old-growth forests of Nothofagus pumilio in Southern ChilePlant Ecology 173(1): 95-106

Other publications

Piper, F.I. & Fajardo, A. 2011. Does Patagonia need dams? An e-letter response to Science article “A craving for hydropower”

Piper, F.I. & Fajardo, A. 2011. Dams and afforestation plans in Chilean Patagonia. Frontiers in Ecology and the Environment 9(2): 96.

Piper, F.I. & Fajardo, A. 2010. Los proyectos hidroeléctricos y los planes de forestación en la Patagonia chilena: Fracaso garantizado? Opinion document.

Fajardo, A. 2010. Reseña de libro “Introducción al análisis espacial de datos en ecología y ciencias ambientales: Métodos y aplicaciones”. Revista Chilena de Historia Natural 83(2): 321-322